This page has been prepared by UNSW Embryology Carnegie Stages, specifically by Dr. M. Hill. Unfortunately the above site has been recently several times unaccessible and because of its usefulness I put up this mirror.
Fly|Worm|Zebrafish|Frog|Chicken|Mouse|Rat|Rabbit|Other Embryos
Carnegie staging is based upon specific developmental features shared by vertebrate embryos. The following graph compares various embryos in relation to Carnegie stages. The data in relation to Days is only an approximation (an average of the range) due to the variability that occurs in normal development. The most common measurement is the crown to rump length (CRL) and the effects of fixation should be considered when making these measurements.
Carnegie is the name of a historical US institute that studies embryology.
Species | Stage | |||||||||||||||
Days |
20 |
22 |
24 |
28 |
30 |
33 |
36 |
40 |
42 |
44 |
48 |
52 |
54 |
55 |
58 |
|
Baboon |
Days |
23 |
25 |
27 |
28 |
29 |
30 |
31 |
33 |
35 |
37 |
39 |
41 |
43 |
45 |
47 |
Rhesus Monkey |
Days |
21 |
22 |
25 |
28 |
29 |
30 |
32 |
34 |
36 |
37 |
38 |
40 |
42 |
44 |
46 |
Marmoset |
Days |
57 |
60 |
64 |
67 |
74 |
||||||||||
Days |
9 |
9.5 |
10 |
10.5 |
11 |
11.5 |
12 |
12.5 |
13 |
13.5 |
14 |
14.5 |
15 |
15.5 |
16 |
|
Rat |
Days |
10.5 |
11 |
11.5 |
12 |
12.5 |
13 |
13.5 |
14 |
14.5 |
15 |
15.5 |
16 |
16.5 |
17 |
17.5 |
Chinese Hamster |
Days |
10 |
10.5 |
11 |
11.5 |
12 |
12.5 |
13 |
13.5 |
14 |
14.5 |
15 |
15.5 |
16 |
16.5 |
17 |
Guinea Pig |
Days |
14.5 |
15 |
15.5 |
17 |
18 |
19 |
20 |
21 |
22 |
23 |
24 |
25 |
26 |
27 |
29 |
Rabbit |
Days |
8 |
8.5 |
9.5 |
10.5 |
11 |
12 |
12.5 |
13.5 |
14 |
14.5 |
15.5 |
16 |
16.5 |
17 |
18 |
Sheep |
Days |
15 |
16 |
17.5 |
18.5 |
19.5 |
20.5 |
22 |
23 |
24.5 |
25.5 |
27.5 |
29.5 |
30 |
33 |
|
Pig |
Days |
14 |
15 |
16 |
17 |
18 |
19 |
20.5 |
21.5 |
23 |
24 |
25.5 |
27.5 |
29 |
30.5 |
32.5 |
Days |
1 |
1.5 |
2 |
2.25 |
2.5 |
3 |
3.25 |
3.75 |
4.75 |
5.5 |
6.25 |
7.25 |
7.75 |
8.5 |
10 |
Source: Butler, H. and Juurlink, B.H.J. An atlas for staging mammalian and chick embryos. CRC Press Inc., Florida (1987) and specific references below.
Species | Source | Reference |
Human | O'Rahilly | Early human development and the chief source of information on staged human embryos. Eur. J. Obstet. Gynec. Reprod. Biol. 9 p273 (1979) |
Baboon | Hendrickx | Embryology of the baboon. University of Chicago Press. |
Rhesus Monkey | Hendrickx and Sawey | Embryology of the rhesus monkey. The Rhesus Monkey, Bourne (ed), Academic Press, NY (1975) |
Marmoset | Phillips | The embryology of the common marmoset. Adv. Anat. Embryol. Cell Biol. 52 p5 (1976) |
Mouse | Theiler | The house mouse. Springer-Verlag, NY (1972) |
Rat | Witschi | Growth. Altman and Dittmer (ed), Fed. Soc. Exp. Biol., Washington (1962) |
Chinese Hamster | Donkelaar | Stages in the prenatal development of the chinese hamster. Anat. Embryol. 156 p1 (1979) |
Guinea Pig | Harman and Prickett | The development of the external form of the guinea-pig between the ages of 11 and 20 days of gestation. Am. J. Anat. 49 p351 (1932) |
Rabbit | Edwards | The external development of the rabbit and rat embryo. Advances in teratology, Vol 3, Woollam (ed), Academic Press, NY (1968) |
Sheep | Bryden | Embryology of the sheep. J. Morphology 138 p169 (1972) |
Pig | Marrable | The embryonic pig. A chronological account. Pitman Medical, London (1971) |
Chicken | Hamburger and Hamilton | A series of normal stages in the development of the chick embryo. J. Morph. 88 p49 (1951) |
Links |
Serial Sections Homepage |
Human Homepage |
Pig Homepage |